Love the weather sealing photo. Wow.
sigh, yet another lens to lust over and I can't afford!
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I shoot wildlife (1Dx) pretty well exclusively, given the time frame you mentioned my suggestion would be the 70-200 f/2.8 V2 with a 1.4x & 2.0x extenders, keeps the weight down and gives you versatility (try to get the V3 extenders, much better optically than the V2 units), I use the 300/400 & 600 V2 lenses as well as the 70-200 and find with all the large Lenses I still use the 70-200 around 50% of the time, with the 70-200 you can zoom in, attach the extenders where required, but with something like the 500 your pretty well stuck if you need to move back, often not an option and by the time you set yourself back up to get the full animal in the shot, the animals long gone, and although the 500f/4 is not a monster, you will note the weight when compared to the 70-200.