The default for long exposure noise reduction is on. Likewise for high ISO noise reduction it is standard. I always switch both to off, so that I have more control and have the maximum frame rate possible. I've never checked to see if noise reduction also affects focus speed/accuracy, but low light focusing is one of the achilles heels of the 7D. One shot is also often more accurate than servo in low light. I have my 5D MkII set to one shot the majority of the time, while on the 7D I have it on AI Servo by default, so that probably increases the difference between the two in low light. The 5D MkII is far superior in terms of accuracy then, so it could be that the less sophisticated focus of the 550D is working to its advantage. Simple is sometimes best,