1DX photo post processed:
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Do you not think three cameras is overkill? Unless you plan on linking them all wirelessly and trigger them that way two would suffice wouldn't it?
Reach is always good, so I would keep the 1D Mark IV and 1D X and sell the rest and buy new glass.
I like Mt Spokane Photography's idea of the new 200-400 lens.
How did you end up with so many cameras anyway?
I really think this is the million dollar (or at least US$ 6,000) question. Once you go past a certain point in length, lenses become super expensive. You can then use extenders as MWP mentioned but then questions re AF and IQ come up. Even a 1.3x crop helps with the reach, and the 1D 4 is about half the price, and you can use extenders on that. If the 1D X had a lot more megapixels you could combat the lack of reach by cropping more, but then you would probably lose out on fps. It is an interesting debate, but the quality of images from the 1D X do look outstanding, and as long as you have lenses that work best (is it groups A-F or something) then you should be on to a winner.