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Should I purchase a 300mm f/2.8 is I for wildlife???

But also, don't f4 lenses focus faster than f5.6?

Not necessarily. It depends on the lens.
With the 500mm F/4 the lens may be moving more mass inside.
Battery power, (other bodies vs the 1D line) seems to matter more for speed on the bigger lenses.

One stop is an 8 to 10 minute advantage in the morning and evening. It may not sound like much but you gain time at the most prime wildlife time.
 
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Not necessarily. It depends on the lens.
With the 500mm F/4 the lens may be moving more mass inside.
Battery power, (other bodies vs the 1D line) seems to matter more for speed on the bigger lenses.

One stop is an 8 to 10 minute advantage in the morning and evening. It may not sound like much but you gain time at the most prime wildlife time.

I meant not in terms of the lens but the AF sensor in the body.
 
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