Ergonomics: grip shape/size, buttons placement, menus and GUIs. When i got my first DSLR, i knew really nothing about the brand wars, i just wanted the best bang for the buck to get started in photography, so i stepped into the shop with the intention of getting a Nikon D3100. Right beside it, there was this shiny and sexy, yet slightly more expensive, EOS 550D. I handled them both, and asked to have them powered on. It was love at first sight with the Canon. 
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