I have no trouble getting excellent wildlife portraits of posing animals and birds with the R (see the puffin above) and the files are beautiful and don't require the cleanup that those from the 7DMKII do. But when I refer to a wildlife camera I need one the can also capture the flying puffin, the attacking leopard, the penguin leaping in the water, the charging bear, and have the focus be in the right spot. The R is more than capable of capturing a leopard sitting in a tree, a penguin standing on the beach, and a bear clamming but these are all static wildlife shots and nice in their own way but I also to shoot a lot of animal behavior which involves more action and I've have not found the R suitable in any way for this. I would certainly be open to any suggestion you have about my settings that could improve this. The picture below was one of a series of rapid shots of a leopard attacking an impala using the 7DMKII. I always want to be prepared for something like this as well as the portraits.I have not had that much trouble getting good to excellent wildlife results from the R using lenses from the 100-400 L IS ii to the 400/2.8 L IS and the 500/4 L IS ii, often with extenders both of necessity and curiosity. But then I started photographing wildlife using thumb drive cameras with manual focus and film. And I got salable results. So I say it is a usable wildlife photography camera.
That said, would I like the guts of a Nikon 850 stuffed into an R body? Yes, I would. That would be my ultimate wildlife and landscape camera.
Catherine
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